Ellen Greene revisits Audrey from 'Little Shop of Horrors' for the film's 40th anniversary

Ellen Greene is revisiting one of her defining roles for the 40th anniversary of “Little Shop of Horrors.”

In conversation, she seamlessly slips in and out of that singular voice — a little breathy, a little squeaky, a little lispy, a little New York — that made her “Little Shop of Horrors” heroine an instant icon. It wasn’t a register she’d planned on hitting at the audition. It just came out of her in the moment, and it stuck.

“I didn’t think about it,” Greene, 75, said in a recent interview with The Associated Press. “It was weird, I mean, I don’t know where half of this stuff came from.”

In her long and robust career on the stage and screen, Audrey is perhaps Greene’s defining role. Her tragicomic shopgirl of skid row belted her way into audiences’ hearts off-Broadway in 1982, and then in Frank Oz’s 1986 film opposite Rick Moranis.

The catchy musical about that flesh-eating plant is never too far in the rearview either. Every few years it seems she’s beckoned for an encore. In 2015, she reprised the role for three sold-out shows, with Jake Gyllenhaal playing Seymour.

Starting this weekend, she'll be making stops around the country for the next few months for 40th anniversary screenings, from Seattle (Aug. 18) to Atlanta (Nov. 13). Performances of “Somewhere That’s Green” have been promised too.

That aching ballad about wanting more, aside from being one of the big showstoppers, holds a special place for Greene. She remembers first listening to it on the unemployment line, waiting to get her check, and quietly singing along.

“I just knew there was an Audrey inside of me waiting to be born,” Greene said.

She carried that feeling to the audition where composer Alan Menken jumped in to play the piano with lyricist Howard Ashman, already her great friend, looking on. When she finished, everyone was in tears.

Getting the role wasn’t any sort of immediate windfall either. When they started at the WPA Theatre in New York, she was being paid $50 a week for a month of rehearsals and two months of shows off-Broadway. But it was a hit. One night, Steve Martin, who would later play Audrey’s abusive dentist boyfriend in the film, even came to see it with Mary Tyler Moore.

On the stage, she performed “Somewhere That's Green” sitting on a trash can. For the film, Ashman wrote the number to be staged like the Leslie Caron and Cyd Charisse numbers they both loved, where a character walks into a room and suddenly they’re in a different outfit for the song.

“You would think I would just have been floored and so excited that he gave me this gift,” she said. “Instead, I call him up and I’m crying my eyes out. And I’m saying, wait for this, ‘Where’s my trash can?’”

Ashman, she said, just laughed and assured her that they’d elicit the same response when the camera pans out to reveal she’s still on skid row. Ironically, this whole exchange happened before she’d even officially been cast in the movie.

The story of how the film was made is long and bumpy. John Landis was originally attached to direct, then, briefly, Martin Scorsese, before it found its way to Oz. Greene’s participation was never a given, either.